What Type Of Animal Uses Beating Flagella To Assist In Filter Feeding?
Chapter fifteen: Diverseness of Animals
Sponges and Cnidarians
Learning Objectives
By the finish of this section, you will be able to:
- Describe the organizational features of the simplest animals
- Depict the organizational features of cnidarians
The kingdom of animals is informally divided into invertebrate animals, those without a backbone, and vertebrate animals, those with a backbone. Although in general we are virtually familiar with vertebrate animals, the vast majority of creature species, near 95 percentage, are invertebrates. Invertebrates include a huge diversity of animals, millions of species in about 32 phyla, which nosotros can but begin to impact hither.
The sponges and the cnidarians stand for the simplest of animals. Sponges appear to stand for an early phase of multicellularity in the brute clade. Although they have specialized cells for particular functions, they lack true tissues in which specialized cells are organized into functional groups. Sponges are like to what might have been the antecedent of animals: colonial, flagellated protists. The cnidarians, or the jellyfish and their kin, are the simplest animal group that displays true tissues, although they possess merely two tissue layers.
Sponges
Animals in subkingdom Parazoa correspond the simplest animals and include the sponges, or phylum Porifera ([Figure 1]). All sponges are aquatic and the bulk of species are marine. Sponges live in intimate contact with water, which plays a role in their feeding, gas exchange, and excretion. Much of the body structure of the sponge is dedicated to moving water through the body then it tin can filter out food, absorb dissolved oxygen, and eliminate wastes.
The torso of the simplest sponges takes the shape of a cylinder with a large central crenel, the spongocoel. H2o enters the spongocoel from numerous pores in the body wall. Water flows out through a big opening chosen the osculum ([Figure 2]). However, sponges exhibit a diversity of torso forms, which vary in the size and branching of the spongocoel, the number of osculi, and where the cells that filter nutrient from the h2o are located.
Sponges consist of an outer layer of flattened cells and an inner layer of cells called choanocytes separated by a jelly-like substance chosen mesohyl. The mesohyl contains embedded amoeboid cells that secrete tiny needles chosen spicules or protein fibers that help give the sponge its structural strength. The cell torso of the choanocyte is embedded in mesohyl but protruding into the spongocoel is a mesh-like neckband surrounding a unmarried flagellum. The beating of flagella from all choanocytes moves water through the sponge. Nutrient particles are trapped in mucus produced by the sieve-like neckband of the choanocytes and are ingested by phagocytosis. This process is chosen intracellular digestion. Amoebocytes have up nutrients repackaged in food vacuoles of the choanocytes and evangelize them to other cells within the sponge.
Physiological Processes in Sponges
Despite their lack of complexity, sponges are conspicuously successful organisms, having persisted on World for more than half a billion years. Lacking a truthful digestive system, sponges depend on the intracellular digestive processes of their choanocytes for their energy intake. The limit of this type of digestion is that food particles must be smaller than individual cells. Gas substitution, circulation, and excretion occur by improvidence between cells and the h2o.
Sponges reproduce both sexually and asexually. Asexual reproduction is either by fragmentation (in which a slice of the sponge breaks off and develops into a new individual), or budding (an outgrowth from the parent that eventually detaches). A type of asexual reproduction found just in freshwater sponges occurs through the germination of gemmules, clusters of cells surrounded by a tough outer layer. Gemmules survive hostile environments and can attach to a substrate and grow into a new sponge.
Sponges are monoecious (or hermaphroditic), meaning 1 individual tin produce both eggs and sperm. Sponges may be sequentially hermaphroditic, producing eggs first and sperm subsequently. Eggs arise from amoebocytes and are retained within the spongocoel, whereas sperm arise from choanocytes and are ejected through the osculum. Sperm carried by water currents fertilize the eggs of other sponges. Early larval development occurs within the sponge, and free-swimming larvae are so released through the osculum. This is the only time that sponges exhibit mobility. Sponges are sessile as adults and spend their lives fastened to a fixed substrate.
Watch this video that demonstrates the feeding of sponges.
Cnidarians
The phylum Cnidaria includes animals that testify radial or biradial symmetry and are diploblastic. Virtually all (about 99 per centum) cnidarians are marine species. Cnidarians have specialized cells known as cnidocytes ("stinging cells") containing organelles called nematocysts. These cells are concentrated effectually the mouth and tentacles of the animal and can immobilize prey with toxins. Nematocysts contain coiled threads that may bear barbs. The outer wall of the cell has a hairlike projection that is sensitive to touch on. When touched, the cells fire the toxin-containing coiled threads that tin can penetrate and stun the predator or casualty (see [Figure 3]).
Cnidarians display two distinct torso plans: polyp or "stalk" and medusa or "bong" ([Effigy 4]). Examples of the polyp form are freshwater species of the genus Hydra; perchance the best-known medusoid animals are the jellies (jellyfish). Polyps are sessile as adults, with a unmarried opening to the digestive system (the mouth) facing up with tentacles surrounding it. Medusae are motile, with the mouth and tentacles hanging from the bell-shaped body. In other cnidarians, both a polyp and medusa class exist, and the life cycle alternates between these forms.
Physiological Processes of Cnidarians
All cnidarians accept ii tissue layers. The outer layer is called the epidermis, whereas the inner layer is called the gastrodermis and lines the digestive cavity. Between these two layers is a non-living, jelly-like mesoglea. There are differentiated cell types in each tissue layer, such as nervus cells, enzyme-secreting cells, and nutrient-absorbing cells, as well every bit intercellular connections between the cells. However, organs and organ systems are not present in this phylum.
The nervous system is primitive, with nerve cells scattered across the torso in a network. The role of the nerve cells is to carry signals from sensory cells and to contractile cells. Groups of cells in the nerve cyberspace form nerve cords that may be essential for more rapid transmission. Cnidarians perform extracellular digestion, with digestion completed past intracellular digestive processes. Food is taken into the gastrovascular cavity, enzymes are secreted into the cavity, and the cells lining the cavity absorb the nutrient products of the extracellular digestive procedure. The gastrovascular cavity has but one opening that serves every bit both a mouth and an anus (an incomplete digestive organization). Like the sponges, Cnidarian cells commutation oxygen, carbon dioxide, and nitrogenous wastes past improvidence betwixt cells in the epidermis and gastrodermis with h2o.
Cnidarian Diversity
The phylum Cnidaria contains about 10,000 described species divided into four classes: Anthozoa, Scyphozoa, Cubozoa, and Hydrozoa.
The class Anthozoa includes all cnidarians that exhibit a sessile polyp body program only; in other words, there is no medusa phase inside their life cycle. Examples include sea anemones, sea pens, and corals, with an estimated number of 6,100 described species. Sea anemones are usually brightly colored and can attain a size of one.8 to 10 cm in bore. These animals are usually cylindrical in shape and are fastened to a substrate. A mouth opening is surrounded by tentacles bearing cnidocytes ([Figure v]).
Scyphozoans include all the jellies and are motile and exclusively marine with virtually 200 described species. The medusa is the ascendant stage in the life bicycle, although there is also a polyp phase. Species range from ii cm in length to the largest scyphozoan species, Cyanea capillata, at 2 g across. Jellies display a characteristic bell-similar body shape ([Figure half dozen]).
Place the life cycle stages of jellies using this video blitheness game from the New England Aquarium.
The class Cubozoa includes jellies that are square in cross-section and and then are known as "box jellyfish." These species may achieve sizes of 15–25 cm. Cubozoans are anatomically similar to the jellyfish. A prominent difference between the two classes is the arrangement of tentacles. Cubozoans have muscular pads called pedalia at the corners of the square bell canopy, with i or more than tentacles attached to each pedalium. In some cases, the digestive system may extend into the pedalia. Cubozoans typically exist in a polyp form that develops from a larva. The polyps may bud to course more polyps and then transform into the medusoid forms.
Picket this video to learn more than about the deadly toxins of the box jellyfish.
Hydrozoa includes virtually 3,500 species, ane virtually of which are marine. Well-nigh species in this class have both polyp and medusa forms in their life bicycle. Many hydrozoans form colonies composed of branches of specialized polyps that share a gastrovascular cavity. Colonies may as well be gratis-floating and contain both medusa and polyp individuals in the colony, equally in the Portuguese Man O'War (Physalia) or By-the-Wind Sailor (Velella). Other species are solitary polyps or lonely medusae. The feature shared by all of these species is that their gonads are derived from epidermal tissue, whereas in all other cnidarians, they are derived from gastrodermal tissue ([Figure 7]ab).
Section Summary
Animals included in phylum Porifera are parazoans and do not possess true tissues. These organisms show a simple organisation. Sponges take multiple prison cell types that are geared toward executing various metabolic functions.
Cnidarians have outer and inner tissue layers sandwiching a noncellular mesoglea. Cnidarians possess a well-formed digestive organisation and carry out extracellular digestion. The cnidocyte is a specialized cell for delivering toxins to prey and predators. Cnidarians have separate sexes. They have a life cycle that involves morphologically distinct forms—medusoid and polypoid—at diverse stages in their life wheel.
Review Questions
The large central opening in the poriferan body is called the _____.
- emmule
- picule
- stia
- osculum
[reveal-answer q="514015″]Prove Answer[/reveal-answer]
[hidden-answer a="514015″]4[/subconscious-answer]
Cnidocytes are institute in _____.
- phylum Porifera
- phylum Nemertea
- phylum Nematoda
- phylum Cnidaria
[reveal-answer q="878659″]Evidence Respond[/reveal-answer]
[hidden-answer a="878659″]iv[/hidden-answer]
Cubozoans are ________.
- polyps
- medusoids
- polymorphs
- sponges
[reveal-respond q="908474″]Prove Answer[/reveal-reply]
[hidden-answer a="908474″]two[/hidden-answer]
Free Response
Describe the feeding mechanism of sponges and identify how information technology is different from other animals.
The sponges describe water carrying food particles into the spongocoel using the beating of flagella in the choanocytes. The food particles are caught by the collar of the choanocyte and brought into the cell by phagocytosis. Digestion of the food particle takes place within the cell. The difference between this and the mechanisms of other animals is that digestion takes place inside cells rather than outside of cells. It means that the organism can feed only on particles smaller than the cells themselves.
Compare the structural differences betwixt Porifera and Cnidaria.
Poriferans practice non possess true tissues, whereas cnidarians do have tissues. Because of this deviation, poriferans exercise not have a nervus net or muscle cells for locomotion, which cnidarians have.
Footnotes
- i "The Hydrozoa Directory," Peter Schuchert, Muséum Genève, last updated November 2012, http://www.ville-ge.ch/mhng/hydrozoa/hydrozoa-directory.htm.
Glossary
- amoebocyte
- an amoeba-similar cell of sponges whose functions include distribution of nutrients to other cells in the sponge
- budding
- a form of asexual reproduction that occurs through the growth of a new organism as a branch on an adult organism that breaks off and becomes contained; found in plants, sponges, cnidarians, and some other invertebrates
- choanocyte
- a cell type unique to sponges with a flagellum surrounded by a collar used to maintain water flow through the sponge, and capture and assimilate food particles
- Cnidaria
- a phylum of animals that are diploblastic and have radial symmetry and stinging cells
- cnidocyte
- a specialized stinging cell found in Cnidaria
- epidermis
- the layer of cells that lines the outer surface of an animal
- extracellular digestion
- a form of digestion, the breakdown of nutrient, which occurs outside of cells with the aid of enzymes released by cells
- fragmentation
- a form of asexual reproduction in which a portion of the body of an organism breaks off and develops into a living independent organism; found in plants, sponges, and another invertebrates
- gastrodermis
- the layer of cells that lines the gastrovascular cavity of cnidarians
- gastrovascular cavity
- the fundamental cavity bounded by the gastrodermis in cnidarians
- gemmule
- a construction produced past asexual reproduction in freshwater sponges that is able to survive harsh atmospheric condition
- intracellular digestion
- the digestion of affair brought into a cell by phagocytosis
- medusa
- a free-floating cnidarian body programme with a oral fissure on the underside and tentacles hanging downwardly from a bell
- mesoglea
- the non-living, gel-like matrix present in between ectoderm and endoderm in cnidarians
- mesohyl
- the collagen-like gel containing suspended cells that perform diverse functions in sponges
- monoecious
- having both sexes in one body, hermaphroditic
- nematocyst
- the harpoon-similar organelle within a cnidocyte with a pointed projectile and toxicant to stun and entangle prey
- osculum
- the large opening in a sponge trunk through which h2o leaves
- polyp
- the stem-like, sessile life class of a cnidarians with oral fissure and tentacles facing upwards, usually sessile merely may be able to glide along a surface
- Porifera
- a phylum of animals with no truthful tissues, but a porous body with a rudimentary endoskeleton
- spicule
- a short sliver or fasten-like structure, in sponges, they are formed of silicon dioxide, calcium carbonate, or protein, and are plant in the mesohyl
- spongocoel
- the fundamental crenel within the body of some sponges
What Type Of Animal Uses Beating Flagella To Assist In Filter Feeding?,
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